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目錄
前言
總論 1
一、國內外研究概況 1
二、形態特徵 3
（一）成蟲 3
（二）幼蟲 23
（三）蛹 26
（四）卵 27
三、分類發展歷史和系統發育關係 27
（一）閻甲總科的系統地位 27
（二）閻甲總科的分類 28
（三）扁圓甲科的系統地位 29
（四）長閻甲科的系統地位 29
（五）閻甲科的分類發展歷史和系統發育關係 30
四、地理分佈 37
（一）屬的分佈類型及其特點 37
（二）種的分佈類型及其特點 39
五、生物學和生態學特性 52
（一）食性和棲息環境 52
（二）世代和發育曆期 55
（三）成蟲的生活習性和行為學 56
（四）天敵 58
六、經濟意義 58
各論 60
閻甲總科Histeroidea Gyllenhal, 1808 60
一、扁圓甲科Sphaeritidae Shuckard, 1839 60
1. 扁圓甲屬Sphaerites Duftschmid, 1805 61
(1) 雙色扁圓甲Sphaerites dimidiatus Jurecek, 1934 61
(2) 黑扁圓甲Sphaerites glabratus (Fabricius, 1792) 63
(3) 翔扁圓甲Sphaerites involatilis Gusakov, 2004 64
(4) 亮扁圓甲Sphaerites nitidus L？bl, 1996 64
(5) 暗扁圓甲Sphaerites opacus L？bl et Háva, 2002 66
(6) 孔扁圓甲Sphaerites perforatus Gusakov, 2017 67
二、長閻甲科Synteliidae Lewis, 1882 67
2. 長閻甲屬Syntelia Westwood, 1864 68
(7) 大衛長閻甲Syntelia davidis Fairmaire, 1889 68
(8) 馬氏長閻甲Syntelia mazuri Zhou, 2003 69
(9) 中華長閻甲Syntelia sinica Zhou, 2003 71
三、閻甲科Histeridae Gyllenhal, 1808 72
（一）細閻甲亞科Niponiinae Fowler, 1912 74
3. 細閻甲屬Niponius Lewis, 1885 74
(10) 溝細閻甲Niponius canalicollis Lewis, 1901 75
(11) 凹細閻甲Niponius impressicollis Lewis, 1885 76
(12) 姬細閻甲Niponius osorioceps Lewis, 1885 77
(13) 山崎細閻甲Niponius yamasakii Miwa, 1934 79
（二）嗜蟻閻甲亞科Chlamydopsinae Bickhardt, 1914 81
4. 奇異閻甲屬Eucurtiopsis Silvestri, 1926 81
(14) 奇異閻甲Eucurtiopsis mirabilis Silvestri, 1926 82
（三）脊閻甲亞科Onthophilinae MacLeay, 1819 82
5. 脊閻甲屬Onthophilus Leach, 1817 83
(15) 黃角脊閻甲Onthophilus flavicornis Lewis, 1884 85
(16) 細脊閻甲Onthophilus foveipennis Lewis, 1885 86
(17) 麗江脊閻甲Onthophilus lijiangensis Zhou et Luo, 2001 88
(18) 原脊閻甲Onthophilus ordinarius Lewis, 1879 90
(19) 粗脊閻甲Onthophilus ostreatus Lewis, 1879 91
(20) 席氏脊閻甲Onthophilus silvae Lewis, 1884 93
(21) 斯氏脊閻甲Onthophilus smetanai Mazur, 1994 94
(22) 瘤脊閻甲Onthophilus tuberculatus Lewis, 1892 95
6. 毛脊閻甲屬Epiechinus Lewis, 1891 95
(23) 多刺毛脊閻甲Epiechinus hispidus (Paykull, 1811) 96
(24) 馬氏毛脊閻甲Epiechinus marseuli Lewis, 1900 97
(25) 南洋毛脊閻甲Epiechinus taprobanae Lewis, 1892 98
(26) 樹毛脊閻甲Epiechinus arboreus (Lewis, 1884) 99
（四）小閻甲亞科Tribalinae Bickhardt, 1914 100
7. 悅閻甲屬Epierus Erichson, 1834 100
(27) 索氏悅閻甲Epierus sauteri Bickhardt, 1913 101
8. 副悅閻甲屬Parepierus Bickhardt, 1913 101
(28) 中國副悅閻甲Parepierus chinensis Zhang et Zhou, 2007 102
(29) 異刺副悅閻甲Parepierus inaequispinus Zhang et Zhou, 2007 104
(30) 劉氏副悅閻甲Parepierus lewisi Bickhardt, 1913 106
(31) 梳刺副悅閻甲Parepierus pectinispinus Zhang et Zhou, 2007 107
9. 小閻甲屬Tribalus Erichson, 1834 109
1) 小閻甲亞屬Tribalus Erichson, 1834 110
(32) 點小閻甲Tribalus (Tribalus) punctillatus Bickhardt, 1913 110
2) 真小閻甲亞屬Eutribalus Bickhardt, 1921 111
(33) 鴿小閻甲Tribalus (Eutribalus) colombius Marseul, 1864 111
(34) 柯氏小閻甲Tribalus (Eutribalus) koenigius Marseul, 1864 113
(35) 歐氏小閻甲Tribalus (Eutribalus) ogieri Marseul, 1864 115
（五）閻甲亞科Histerinae Gyllenhal, 1808 115
一）突胸閻甲族Exosternini Bickhardt, 1914 116
10. 短臀閻甲屬Anaglymma Lewis, 1894 117
(36) 環短臀閻甲Anaglymma circularis (Marseul, 1864) 117
11. 圓臀閻甲屬Notodoma Lacordaire, 1854 119
(37) 蕈圓臀閻甲Notodoma fungorum Lewis, 1884 120
12. 凹背閻甲屬Epitoxus Lewis, 1900 122
(38) 亞洲凹背閻甲Epitoxus asiaticus Vienna, 1986 123
(39) 布拉凹背閻甲Epitoxus bullatus (Marseul, 1870) 123
13. 長臀閻甲屬Cypturus Erichson, 1834 124
(40) 埃長臀閻甲Cypturus aenescens Erichson, 1834 125
14. 緣尾閻甲屬Paratropus Gerstaecker, 1867 127
(41) 坎達拉緣尾閻甲Paratropus khandalensis Kanaar, 1997 127
二）扁閻甲族Hololeptini Hope, 1840 129
15. 扁閻甲屬Hololepta Paykull, 1811 130
3) 扁閻甲亞屬Hololepta Paykull, 1811 131
(42) 烏蘇裡扁閻甲Hololepta (Hololepta) amurensis Reitter, 1879 131
(43) 鮑氏扁閻甲Hololepta (Hololepta) baulnyi Marseul, 1857 133
(44) 扁閻甲Hololepta (Hololepta) depressa Lewis, 1884 135
(45) 長扁閻甲Hololepta (Hololepta) elongata Erichson, 1834 137
(46) 費氏扁閻甲Hololepta (Hololepta) feae Lewis, 1892 138
(47) 希氏扁閻甲Hololepta (Hololepta) higoniae Lewis, 1894 140
(48) 印度扁閻甲Hololepta (Hololepta) indica Erichson, 1834 141
(49) 亮扁閻甲Hololepta (Hololepta) laevigata Guérin-Ménéville, 1833 143
(50) 尼泊爾扁閻甲Hololepta (Hololepta) nepalensis Lewis, 1910 145
(51) 鈍扁閻甲Hololepta (Hololepta) obtusipes Marseul, 1864 145
(52) 平扁閻甲Hololepta (Hololepta) plana (Sulzer, 1776) 147
三）方閻甲族Platysomatini Bickhardt, 1914 148
16. 巨顎閻甲屬Megagnathos Penati et Zhang, 2009 150
(53) 巨顎閻甲Megagnathos lagardei Vienna et Ratto, 2015 151
17. 溝顎閻甲屬Silinus Lewis, 1907 151
(54) 溝顎閻甲Silinus procerus (Lewis, 1911) 151
18. 長卵閻甲屬Platylister Lewis, 1892 153
4) 長卵閻甲亞屬Platylister Lewis, 1892 154
(55) 黑長卵閻甲Platylister (Platylister) atratus (Erichson, 1834) 154
(56) 緬甸長卵閻甲Platylister (Platylister) birmanus (Marseul, 1861) 156
(57) 柬埔寨長卵閻甲Platylister (Platylister) cambodjensis (Marseul, 1864) 158
(58) 霍氏長卵閻甲Platylister (Platylister) horni (Bickhardt, 1913) 161
(59) 平氏長卵閻甲Platylister (Platylister) pini (Lewis, 1884) 162
(60) 縫長卵閻甲Platylister (Platylister) suturalis (Lewis, 1888) 164
(61) 凱氏長卵閻甲Platylister (Platylister) cathayi Lewis, 1900 165
5) 平尾閻甲亞屬Popinus Mazur, 1999 166
(62) 孔氏長卵閻甲Platylister (Popinus) confucii (Marseul, 1857) 166
(63) 篩臀長卵閻甲Platylister (Popinus) dahdah (Marseul, 1861) 168
(64) 光亮長卵閻甲Platylister (Popinus) lucillus (Lewis, 1891) 169
(65) 獨長卵閻甲Platylister (Popinus) unicus (Bickhardt, 1912) 171
19. 闊咽閻甲屬Apobletes Marseul, 1860 173
(66) 緣闊咽閻甲Apobletes marginicollis Lewis, 1888 173
(67) 紹氏闊咽閻甲Apobletes schaumei Marseul, 1860 175
20. 方閻甲屬Platysoma Leach, 1817 178
6) 方閻甲亞屬Platysoma Leach, 1817 179
(68) 貝氏方閻甲Platysoma (Platysoma) beybienkoi Kryzhanovskij, 1972 179
(69) 短線方閻甲Platysoma (Platysoma) brevistriatum Lewis, 1888 179
(70) 中國方閻甲Platysoma (Platysoma) chinense Lewis, 1894 181
(71) 平方閻甲Platysoma (Platysoma) deplanatum (Gyllenhal, 1808) 183
(72) 達氏方閻甲Platysoma (Platysoma) dufali Marseul, 1864 185
(73) 重方閻甲Platysoma (Platysoma) gemellun (Cooman, 1929) 187
(74) 朝鮮方閻甲Platysoma (Platysoma) koreanum Mazur, 1999 189
(75) 滑方閻甲Platysoma (Platysoma) rasile Lewis, 1884 189
(76) 裂方閻甲Platysoma (Platysoma) rimarium Erichson, 1834 189
(77) 四川方閻甲Platysoma (Platysoma) sichuanum Mazur, 2007 189
(78) 井上方閻甲Platysoma (Platysoma) takehikoi ？hara, 1986 191
7) 長方閻甲亞屬Cylister Cooman, 1941 192
(79) 狹方閻甲Platysoma (Cylister) angustatum (Hoffmann, 1803) 193
(80) 長方閻甲Platysoma (Cylister) elongatum (Thunberg, 1787) 195
(81) 線方閻甲Platysoma (Cylister) lineare Erichson, 1834 197
(82) 細方閻甲Platysoma (Cylister) lineicolle Marseul, 1873 199
(83) 雲南方閻甲Platysoma (Cylister) yunnanum (Kryzhanovskij, 1972) 201
21. 近方閻甲屬Niposoma Mazur, 1999 201
(84) 劉氏近方閻甲Niposoma lewisi (Marseul, 1873) 202
(85) 申氏近方閻甲Niposoma schenklingi (Bickhardt, 1913) 204
(86) 斯氏近方閻甲Niposoma stackelbergi (Kryzhanovskij, 1976) 205
(87) 臺灣近方閻甲Niposoma taiwanum (Hisamatsu, 1965) 206
22. 卡那閻甲屬Kanaarister Mazur, 1999 207
(88) 阿薩姆卡那閻甲Kanaarister assamensis (Lewis, 1900) 207
(89) 隱卡那閻甲Kanaarister celatum (Lewis, 1884) 208
(90) 庫氏卡那閻甲Kanaarister coomani (Thérond, 1955) comb. nov. 209
23. 似真卵閻甲屬Eurosomides Newton, 2015 211
(91) 微似真卵閻甲Eurosomides minor (Rossi, 1792) 211
24. 溝尾閻甲屬Liopygus Lewis, 1891 213
(92) 溝尾閻甲Liopygus andrewesi Lewis, 1906 214
25. 短卵閻甲屬Eblisia Lewis, 1889 216
8) 短卵閻甲亞屬Eblisia Lewis, 1889 217
(93) 田舍短卵閻甲Eblisia (Eblisia) pagana Lewis, 1902 217
(94) 小短卵閻甲Eblisia (Eblisia) pygmaea (Bickhardt, 1913) 218
(95) 索氏短卵閻甲Eblisia (Eblisia) sauteri (Bickhardt, 1912) 219
(96) 蘇門答臘短卵閻甲Eblisia (Eblisia) sumatrana (Bickhardt, 1912) 221
9) 點尾閻甲亞屬Chronus Lewis, 1914 222
(97) 凱西短卵閻甲Eblisia (Chronus) calceata (Cooman, 1931) 222
26. 真卵閻甲屬Eurylister Bickhardt, 1920 224
(98) 薩氏真卵閻甲Eurylister satzumae (Lewis, 1899) 225
(99) 樹真卵閻甲Eurylister silvestre (Schmidt, 1897) 226
27. 直溝閻甲屬Mendelius Lewis, 1908 228
(100) 細直溝閻甲Mendelius tenuipes (Lewis, 1905) 228
28. 完額閻甲屬Aulacosternus Marseul, 1853 229
(101) 完額閻甲Aulacosternus zelandicus Marseul, 1853 229
29. 大閻甲屬Plaesius Erichson, 1834 231
10) 大閻甲亞屬Plaesius Erichson, 1834 232
(102) 爪哇大閻甲Plaesius (Plaesius) javanus Erichson, 1834 232
(103) 莫氏大閻甲Plaesius (Plaesius) mohouti Lewis, 1879 234
11) 異胸閻甲亞屬Hyposolenus Lewis, 1907 236
(104) 孟加拉大閻甲Plaesius (Hyposolenus) bengalensis Lewis, 1906 236
四）斜臀閻甲族Omalodini Kryzhanovskij, 1972 238
30. 脊額閻甲屬Lewisister Bickhardt, 1912 239
(105) 脊額閻甲Lewisister excellens Bickhardt, 1912 239
五）閻甲族Histerini Gyllenhal, 1808 241
31. 歧閻甲屬Margarinotus Marseul, 1853 242
12) 光折閻甲亞屬Ptomister Houlbert et Monnot, 1923 244
(106) 阿葛歧閻甲Margarinotus (Ptomister) agnatus (Lewis, 1884) 245
(107) 阿若歧閻甲Margarinotus (Ptomister) arrosor (Bickhardt, 1920) 247
(108) 康夫歧閻甲Margarinotus (Ptomister) babai ？hara, 1999 247
(109) 博氏歧閻甲Margarinotus (Ptomister) boleti (Lewis, 1884) 249
(110) 屍生歧閻甲Margarinotus (Ptomister) cadavericola (Bickhardt, 1920) 251
(111) 海拉爾歧閻甲Margarinotus (Ptomister) hailar Wenzel, 1944 253
(112) 隱歧閻甲Margarinotus (Ptomister) incognitus (Marseul, 1854) 254
(113) 科氏歧閻甲Margarinotus (Ptomister) koltzei (Schmidt, 1889) 256
(114) 多齒歧閻甲Margarinotus (Ptomister) multidens (Schmidt, 1889) 257
(115) 大澤歧閻甲Margarinotus (Ptomister) osawai ？hara, 1999 259
(116) 理氏歧閻甲Margarinotus (Ptomister) reichardti Kryzhanovskij et Reichardt, 1976 261
(117) 紋歧閻甲指名亞種Margarinotus (Ptomister) striola striola (Sahlberg, 1819) 263
(118) 縫連歧閻甲Margarinotus (Ptomister) sutus (Lewis, 1884) 265
(119) 三線歧閻甲Margarinotus (Ptomister) tristriatus Wenzel, 1944 266
(120) 溫氏歧閻甲Margarinotus (Ptomister) wenzelisnus Kryzhanovskij et Reichardt, 1976 266
(121) 魏氏歧閻甲Margarinotus (Ptomister) weymarni Wenzel, 1944 268
13) 毛折閻甲亞屬Eucalohister Reitter, 1909 270
(122) 雙斑歧閻甲Margarinotus (Eucalohister) bipustulatus (Schrank, 1781) 270
(123) 美斑歧閻甲Margarinotus (Eucalohister) gratiosus (Mannerheim, 1852) 272
14) 寬脛閻甲亞屬Stenister Reichardt, 1926 274
(124) 暗歧閻甲Margarinotus (Stenister) obscurus (Kugelann, 1792) 274
15) 端線閻甲亞屬Paralister Bickhardt, 1917 277
(125) 柯氏歧閻甲Margarinotus (Paralister) koenigi (Schmidt, 1888) 277
(126) 中亞歧閻甲Margarinotus (Paralister) laevifossa (Schmidt, 1890) 279
(127) 長圓歧閻甲Margarinotus (Paralister) oblongulus (Schmidt, 1892) 281
(128) 周歧閻甲Margarinotus (Paralister) periphaerus Mazur, 2003 282
(129) 暗紅歧閻甲Margarinotus (Paralister) purpurascens (Herbst, 1792) 284
16) 普脛閻甲亞屬Grammostethus Lewis, 1906 286
(130) 緬甸歧閻甲Margarinotus (Grammostethus) birmanus Lundgren, 1991 287
(131) 臺灣歧閻甲Margarinotus (Grammostethus) formosanus Ohara, 1999 287
(132) 脆歧閻甲Margarinotus (Grammostethus) fragosus (Lewis, 1892) 289
(133) 勤歧閻甲Margarinotus (Grammostethus) impiger (Lewis, 1905) 291
(134) 日本歧閻甲Margarinotus (Grammostethus) niponicus (Lewis, 1895) 292
(135) 海西歧閻甲Margarinotus (Grammostethus) occidentalis (Lewis, 1885) 294
(136) 施氏歧閻甲Margarinotus (Grammostethus) schneideri Kapler, 1996 296
(137) 糞歧閻甲Margarinotus (Grammostethus) stercoriger (Marseul, 1880) 297
(138) 台島歧閻甲Margarinotus (Grammostethus) taiwanus Mazur, 2008 299
17) 縫歧閻甲亞屬Asterister Desbordes, 1920 299
(139) 完縫歧閻甲Margarinotus (Asterister) curvicollis (Bickhardt, 1913) 299
32. 突唇閻甲屬Pachylister Lewis, 1904 301
18) 突唇閻甲亞屬Pachylister Lewis, 1904 302
(140) 斯里蘭卡突唇閻甲寬臀亞種Pachylister (Pachylister) ceylanus pygidialis Lewis, 1906 302
(141) 泥突唇閻甲Pachylister (Pachylister) lutarius (Erichson, 1834) 304
19) 溝唇閻甲亞屬Sulcignathos Mazur, 2010 306
(142) 拙突唇閻甲Pachylister (Sulcignathos) scaevola (Erichson, 1834) 306
33. 完折閻甲屬Nasaltus Mazur et W？grzynowicz, 2008 306
(143) 中國完折閻甲Nasaltus chinensis (Quensel, 1806) 307
(144) 東方完折閻甲Nasaltus orientalis (Paykull, 1811) 310
34. 閻甲屬Hister Linnaeus, 1758 312
(145) 比斯閻甲Hister bissexstriatus Fabricius, 1801 314
(146) 同色閻甲Hister concolor Lewis, 1884 316
(147) 康吉閻甲Hister congener Schmidt, 1885 317
(148) 遙閻甲Hister distans Fischer de Waldheim, 1823 320
(149) 幻異閻甲Hister inexspectatus Desbordes, 1923 322
(150) 日本閻甲Hister japonicus Marseul, 1854 322
(151) 爪哇閻甲Hister javanicus Paykull, 1811 324
(152) 馬氏閻甲Hister mazuri Kapler, 1997 326
(153) 巨爪閻甲Hister megalonyx Reichardt, 1922 328
(154) 普然閻甲Hister pransus Lewis, 1892 330
(155) 點足閻甲Hister punctifemur Mazur, 2010 332
(156) 糙閻甲東部亞種Hister salebrosus subsolanus (Newton, 1991) 332
(157) 謝氏閻甲Hister sedakovii Marseul, 1861 334
(158) 上海閻甲Hister shanghaicus Marseul, 1861 337
(159) 西伯利亞閻甲Hister sibiricus Marseul, 1854 339
(160) 簡胸閻甲Hister simplicisternus Lewis, 1879 341
(161) 斯普閻甲Hister spurius Marseul, 1861 342
(162) 西藏閻甲Hister thibetanus Marseul, 1857 344
(163) 單色閻甲累氏亞種Hister unicolor leonhardi Bickhardt, 1910 346
35. 分閻甲屬Merohister Reitter, 1909 348
(164) 吉氏分閻甲Merohister jekeli (Marseul, 1857) 349
36. 新植閻甲屬Neosantalus Kryzhanovskij, 1972 351
(165) 新植閻甲Neosantalus latitibius (Marseul, 1861) 352
37. 三線閻甲屬Eudiplister Reitter, 1909 355
(166) 米氏三線閻甲Eudiplister muelleri Kryzhanovskij et Reichardt, 1976 355
(167) 平坦三線閻甲Eudiplister planulus (Ménétries, 1848) 356
38. 清亮閻甲屬Atholus Thomson, 1859 358
(168) 奇額清亮閻甲Atholus bifrons (Marseul, 1854) 359
(169) 雙斑清亮閻甲Atholus bimaculatus (Linnaeus, 1758) 360
(170) 青色清亮閻甲Atholus coelestis (Marseul, 1857) 360
(171) 緣清亮閻甲Atholus confinis (Erichson, 1834) 362
(172) 窩胸清亮閻甲Atholus depistor (Marseul, 1873) 362
(173) 十二紋清亮閻甲十四紋亞種Atholus duodecimstriatus quatuordecimstriatus (Gyllenhal, 1808) 365
(174) 勒清亮閻甲Atholus levis Mazur, 2015 367
(175) 菲律賓清亮閻甲Atholus philippinensis (Marseul, 1854) 367
(176) 皮瑞清亮閻甲Atholus pirithous (Marseul, 1873) 369
(177) 紋尾清亮閻甲Atholus striatipennis (Lewis, 1892) 372
(178) 扭清亮閻甲Atholus torquatus (Marseul, 1854) 373
39. 毛腹閻甲屬Asiaster Cooman, 1948 376
(179) 卡卡毛腹閻甲Asiaster calcator Cooman, 1948 376
(180) 庫氏毛腹閻甲Asiaster cooteri Kapler, 1999 378
(181) 赫氏毛腹閻甲Asiaster hlavaci Lackner, 2004 379
（六）伴閻甲亞科Hetaeriinae Marseul, 1857 380
40. 伴閻甲屬Hetaerius Erichson, 1834 381
(182) 悅伴閻甲Hetaerius optatus Lewis, 1884 381
（七）卵閻甲亞科Dendrophilinae Reitter, 1909 383
六）卵閻甲族Dendrophilini Reitter, 1909 384
41. 卵閻甲屬Dendrophilus Leach, 1817 384
20) 卵閻甲亞屬Dendrophilus Leach, 1817 385
(183) 寬卵閻甲Dendrophilus (Dendrophilus) xavieri Marseul, 1873 385
21) 擬卵閻甲亞屬Dendrophilopsis Schmidt, 1890 387
(184) 展卵閻甲Dendrophilus (Dendrophilopsis) proditor (Reichardt, 1925) 387
七）小齒閻甲族Bacaniini Kryzhanovskij, 1976 389
42. 穆勒閻甲屬Mullerister Cooman, 1936 389
(185) 日本穆勒閻甲Mullerister niponicus (Lewis, 1879) 389
(186) 東京灣穆勒閻甲Mullerister tonkinensis (Cooman, 1936) 391
43. 小齒閻甲屬Bacanius LeConte, 1853 391
22) 小齒閻甲亞屬Bacanius LeConte, 1853 392
(187) 科氏小齒閻甲Bacanius (Bacanius) collettei Gomy, 1999 392
(188) 卡氏小齒閻甲Bacanius (Bacanius) kapleri Gomy, 1999 394
(189) 天皇小齒閻甲Bacanius (Bacanius) mikado (Lewis, 1892) 395
44. 顯臀閻甲屬Abraeomorphus Reitter, 1886 397
(190) 臺灣顯臀閻甲Abraeomorphus formosanus (Hisamatsu, 1965) 397
八）麗尾閻甲族Paromalini Reitter, 1909 399
45. 簇點閻甲屬Coomanister Kryzhanovskij, 1972 400
(191) 斯氏簇點閻甲Coomanister scolyti Mazur, 2007 400
46. 勻點閻甲屬Carcinops Marseul, 1855 402
23) 勻點閻甲亞屬Carcinops Marseul, 1855 403
(192) 貝氏勻點閻甲Carcinops (Carcinops) penatii Zhang et Zhou, 2007 403
(193) 小勻點閻甲Carcinops (Carcinops) pumilio (Erichson, 1834) 405
(194) 中華勻點閻甲Carcinops (Carcinops) sinensis Lewis, 1909 408
(195) 隱勻點閻甲Carcinops (Carcinops) troglodytes (Paykull, 1811) 409
47. 胸線閻甲屬Diplostix Bickhardt, 1921 411
24) 胸線閻甲亞屬Diplostix Bickhardt, 1921 412
(196) 卡倫胸線閻甲Diplostix (Diplostix) karenensis (Lewis, 1891) 412
(197) 異胸線閻甲Diplostix (Diplostix) vicaria (Cooman, 1935) 412
48. 厚閻甲屬Pachylomalus Schmidt, 1897 415
25) 額線閻甲亞屬Canidius Cooman, 1941 415
(198) 缺線厚閻甲Pachylomalus (Canidius) deficiens Cooman, 1933 416
(199) 肌厚閻甲Pachylomalus (Canidius) musculus (Marseul, 1873) 419
49. 隱閻甲屬Cryptomalus Mazur, 1993 421
(200) 名豪隱閻甲Cryptomalus mingh Mazur, 2007 421
50. 平閻甲屬Platylomalus Cooman, 1948 423
(201) 斯里蘭卡平閻甲Platylomalus ceylanicus (Motschulsky, 1863) 424
(202) 折平閻甲Platylomalus inflexus Zhang et Zhou, 2007 426
(203) 門第平閻甲Platylomalus mendicus (Lewis, 1892) 428
(204) 日本平閻甲Platylomalus niponensis (Lewis, 1899) 431
(205) 洋平閻甲Platylomalus oceanitis (Marseul, 1855) 433
(206) 索氏平閻甲Platylomalus sauteri (Bickhardt, 1912) 435
(207) 胸線平閻甲Platylomalus submetallicus (Lewis, 1892) 436
(208) 東京灣平閻甲Platylomalus tonkinensis (Cooman, 1937) 439
(209) 旅平閻甲Platylomalus viaticus (Lewis, 1892) 441
51. 簡額閻甲屬Eulomalus Cooman, 1937 443
(210) 安普簡額閻甲Eulomalus amplipes Cooman, 1937 443
(211) 龍目簡額閻甲Eulomalus lombokanus Cooman, 1937 445
(212) 普普簡額閻甲Eulomalus pupulus Cooman, 1937 446
(213) 皺簡額閻甲Eulomalus rugosus Zhang et Zhou, 2007 448
(214) 賽氏簡額閻甲Eulomalus seitzi Cooman, 1941 450
(215) 塔蒂簡額閻甲Eulomalus tardipes (Lewis, 1892) 452
(216) 蠕尾簡額閻甲Eulomalus vermicipygus Cooman, 1937 454
52. 麗尾閻甲屬Paromalus Erichson, 1834 456
26) 麗尾閻甲亞屬Paromalus Erichson, 1834 456
(217) 銳角麗尾閻甲Paromalus (Paromalus) acutangulus Zhang et Zhou, 2007 457
(218) 菱麗尾閻甲Paromalus (Paromalus) parallelepipedus (Herbst, 1792) 459
(219) 皮克麗尾閻甲Paromalus (Paromalus) picturatus Kapler, 1999 461
(220) 西藏麗尾閻甲Paromalus (Paromalus) tibetanus Zhang et Zhou, 2007 463
(221) 春麗尾閻甲Paromalus (Paromalus) vernalis Lewis, 1892 465
（八）球閻甲亞科Abraeinae MacLeay, 1819 467
九）球閻甲族Abraeini MacLeay, 1819 468
53. 刺球閻甲屬Chaetabraeus Portevin, 1929 468
27) 刺球閻甲亞屬Chaetabraeus Portevin, 1929 469
(222) 邦刺球閻甲Chaetabraeus (Chaetabraeus) bonzicus (Marseul, 1873) 469
(223) 合刺球閻甲Chaetabraeus (Chaetabraeus) cohaeres (Lewis, 1898) 471
(224) 粒刺球閻甲Chaetabraeus (Chaetabraeus) granosus (Motschulsky, 1863) 472
(225) 東方刺球閻甲Chaetabraeus (Chaetabraeus) orientalis (Lewis, 1907) 472
28) 馬儒閻甲亞屬Mazureus Gomy, 1991 474
(226) 等刺球閻甲Chaetabraeus (Mazureus) paria (Marseul, 1856) 474
十）斷胸閻甲族Plegaderini Portevin, 1929 475
54. 斷胸閻甲屬Plegaderus Erichson, 1834 475
29) 斷胸閻甲亞屬Plegaderus Erichson, 1834 476
(227) 斷胸閻甲Plegaderus (Plegaderus) vulneratus (Panzer, 1797) 476
十一）異跗閻甲族Acritini Wenzel, 1944 478
55. 阿拜閻甲屬Abaeletes Cooman, 1940 478
(228) 派氏阿拜閻甲Abaeletes perroti (Cooman, 1940) 478
56. 異跗閻甲屬Acritus LeConte, 1853 478
30) 密紋閻甲亞屬Pycnacritus Casey, 1916 480
(229) 白水隆異跗閻甲Acritus (Pycnacritus) shirozui Hisamatsu, 1965 480
31) 異跗閻甲亞屬Acritus LeConte, 1853 481
(230) 庫氏異跗閻甲Acritus (Acritus) cooteri Gomy, 1999 481
(231) 駒井異跗閻甲Acritus (Acritus) komai Lewis, 1879 482
(232) 嗜草異跗閻甲Acritus (Acritus) pascuarum Cooman, 1947 484
(233) 梳異跗閻甲Acritus (Acritus) pectinatus Cooman, 1932 485
(234) 瘤胸異跗閻甲Acritus (Acritus) tuberisternus Cooman, 1932 487
十二）條閻甲族Teretriini Bickhardt, 1916 487
57. 條閻甲屬Teretrius Erichson, 1834 488
32) 新條閻甲亞屬Neoteretrius Kryzhanovskij et Reichardt, 1976 488
(235) 臺灣條閻甲Teretrius (Neoteretrius) formosus (Lewis, 1915) 488
(236) 柴田條閻甲Teretrius (Neoteretrius) shibatai ？hara, 2008 489
33) 條閻甲亞屬Teretrius Erichson, 1834 489
(237) 太極條閻甲Teretrius (Teretrius) taichii ？hara, 2008 489
（九）腐閻甲亞科Saprininae Blanchard, 1845 489
58. 禿額閻甲屬Gnathoncus Jacquelin-Duval, 1858 491
(238) 短胸禿額閻甲Gnathoncus brevisternus Lewis, 1907 492
(239) 迪斯禿額閻甲長縫亞種Gnathoncus disjunctus suturifer Reitter, 1896 492
(240) 克氏禿額閻甲Gnathoncus kiritshenkoi Reichardt, 1930 494
(241) 小齒禿額閻甲Gnathoncus nannetensis (Marseul, 1862) 496
(242) 煙禿額閻甲Gnathoncus nidorum Stockmann, 1957 498
(243) 波氏禿額閻甲Gnathoncus potanini Reitter, 1896 498
(244) 圓禿額閻甲Gnathoncus rotundatus (Kugelann, 1792) 499
(245) 半緣禿額閻甲Gnathoncus semimarginatus Bickhardt, 1920 501
59. 腐閻甲屬Saprinus Erichson, 1834 502
34) 狹胸閻甲亞屬Phaonius Reichardt, 1941 504
(246) 法老腐閻甲Saprinus (Phaonius) pharao Marseul, 1855 504
35) 腐閻甲亞屬Saprinus Erichson, 1834 507
(247) 阿登腐閻甲Saprinus (Saprinus) addendus Dahlgren, 1964 507
(248) 埃腐閻甲Saprinus (Saprinus) aeneolus Marseul, 1870 507
(249) 雙斑腐閻甲Saprinus (Saprinus) biguttatus (Steven, 1806) 510
(250) 圓斑腐閻甲Saprinus (Saprinus) bimaculatus Dahlgren, 1964 512
(251) 藍斑腐閻甲指名亞種Saprinus (Saprinus) caerulescens caerulescens (Hoffmann, 1803) 514
(252) 藍斑腐閻甲點胸亞種Saprinus (Saprinus) caerulescens punctisternus Lewis, 1900 515
(253) 中央腐閻甲Saprinus (Saprinus) centralis Dahlgren, 1971 518
(254) 金澤腐閻甲Saprinus (Saprinus) chalcites (Illiger, 1807) 518
(255) 齊腐閻甲Saprinus (Saprinus) concinnus (Gebler, 1830) 521
(256) 緩腐閻甲Saprinus (Saprinus) dussaulti Marseul, 1870 523
(257) 麗斑腐閻甲Saprinus (Saprinus) flexuosofasciatus Motschulsky, 1845 524
(258) 額紋腐閻甲Saprinus (Saprinus) frontistrius Marseul, 1855 526
(259) 寒鴉腐閻甲Saprinus (Saprinus) graculus Reichardt, 1930 528
(260) 哈氏腐閻甲Saprinus (Saprinus) havajirii Kapler, 2000 530
(261) 喜馬拉雅腐閻甲Saprinus (Saprinus) himalajicus Dahlgren, 1971 531
(262) 汙腐閻甲Saprinus (Saprinus) immundus (Gyllenhal, 1827) 531
(263) 銅澤腐閻甲Saprinus (Saprinus) intractabilis Reichardt, 1930 533
(264) 日本腐閻甲Saprinus (Saprinus) niponicus Dahlgren, 1962 535
(265) 麗鞘腐閻甲Saprinus (Saprinus) optabilis Marseul, 1855 538
(266) 扭斑腐閻甲Saprinus (Saprinus) ornatus Erichson, 1834 540
(267) 派腐閻甲Saprinus (Saprinus) pecuinus Marseul, 1855 543
(268) 平盾腐閻甲Saprinus (Saprinus) planiusculus Motschulsky, 1849 545
(269) 側斑腐閻甲Saprinus (Saprinus) quadriguttatus (Fabricius, 1798) 548
(270) 謝氏腐閻甲Saprinus (Saprinus) sedakovii Motschulsky, 1860 550
(271) 半線腐閻甲Saprinus (Saprinus) semistriatus (Scriba, 1790) 553
(272) 斯博腐閻甲Saprinus (Saprinus) spernax Marseul, 1862 555
(273) 燦腐閻甲Saprinus (Saprinus) splendens (Paykull, 1811) 556
(274) 斯達腐閻甲Saprinus (Saprinus) sternifossa Müller, 1937 558
(275) 淡黑腐閻甲Saprinus (Saprinus) subcoerulus Thérond, 1978 561
(276) 細紋腐閻甲歐亞亞種Saprinus (Saprinus) tenuistrius sparsutus Solskiy, 1876 561
60. 半腐閻甲屬Hemisaprinus Kryzhanovskij, 1976 564
(277) 墨半腐閻甲Hemisaprinus subvirescens (Ménétries, 1832) 564
61. 連窩閻甲屬Chalcionellus Reichardt, 1932 566
(278) 阿莫連窩閻甲Chalcionellus amoenus (Erichson, 1834) 567
(279) 布氏連窩閻甲指名亞種Chalcionellus blanchii blanchii (Marseul, 1855) 569
(280) 布氏連窩閻甲陶芮亞種Chalcionellus blanchii tauricus (Marseul, 1862) 571
(281) 西伯利亞連窩閻甲Chalcionellus sibiricus Dahlgren, 1969 571
(282) 土耳其連窩閻甲Chalcionellus turcicus (Marseul, 1857) 571
62. 多紋閻甲屬Pholioxenus Reichardt, 1932 573
(283) 多紋閻甲Pholioxenus orion Reichardt, 1932 574
63. 連線閻甲屬Hypocacculus Bickhardt, 1914 574
36) 連線閻甲亞屬Hypocacculus Bickhardt, 1914 575
(284) 光額連線閻甲Hypocacculus (Hypocacculus) spretulus (Erichson, 1834) 575
64. 皺額閻甲屬Hypocaccus Thomson, 1867 577
37) 皺額閻甲亞屬Hypocaccus Thomson, 1867 578
(285) 巴西皺額閻甲Hypocaccus (Hypocaccus) brasiliensis (Paykull, 1811) 578
(286) 多芮皺額閻甲Hypocaccus (Hypocaccus) dauricus Reichardt, 1930 578
(287) 皺額閻甲Hypocaccus (Hypocaccus) rugifrons (Paykull, 1798) 580
(288) 辛氏皺額閻甲Hypocaccus (Hypocaccus) sinae (Marseul, 1862) 583
38) 滑背閻甲亞屬Baeckmanniolus Reichardt, 1926 585
(289) 變線皺額閻甲指名亞種Hypocaccus (Baeckmanniolus) varians varians (Schmidt, 1890) 585
(290) 變線皺額閻甲大陸亞種Hypocaccus (Baeckmanniolus) varians continentalis (Reichardt, 1941) 587
39) 粗額閻甲亞屬Nessus Reichardt, 1932 588
(291) 潔皺額閻甲Hypocaccus (Nessus) asticus Lewis, 1911 588
(292) 巴魯皺額閻甲Hypocaccus (Nessus) balux Reichardt, 1932 590
(293) 沙漠皺額閻甲Hypocaccus (Nessus) eremobius Reichardt, 1932 591
(294) 蒙古皺額閻甲Hypocaccus (Nessus) mongolicus Reichardt, 1932 591
(295) 虎皺額閻甲指名亞種Hypocaccus (Nessus) tigris tigris (Marseul, 1862) 592
65. 稀閻甲屬Reichardtiolus Kryzhanovskij, 1959 593
(296) 杜芮稀閻甲Reichardtiolus duriculus (Reitter, 1904) 593
66. 新厚閻甲屬Eopachylopus Reichardt, 1926 594
(297) 溪新厚閻甲Eopachylopus ripae (Lewis, 1885) 594
（十）柱閻甲亞科Trypeticinae Bickhardt, 1913 595
67. 柱閻甲屬Trypeticus Marseul, 1864 595
(298) 溝額柱閻甲Trypeticus canalifrons Bickhardt, 1913 597
(299) 裂吻柱閻甲Trypeticus fissirostrum Zhang et Zhou, 2007 599
(300) 森柱閻甲Trypeticus nemorivagus Lewis, 1892 601
(301) 索氏柱閻甲Trypeticus sauteri Bickhardt, 1913 602
(302) 獵柱閻甲Trypeticus venator (Lewis, 1884) 603
(303) 雲南柱閻甲Trypeticus yunnanensis Zhang et Zhou, 2007 605
參考文獻 609
英文摘要 646
中名索引 680
學名索引 687
《中國動物志》已出版書目 703
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